The islands of Macaronesia are volcanic in origin, and are thought to be the product of several geologic hotspots.

The climate of the Macaronesian islands ranges from subtropical to tropical. The Portuguese archipelagos of the Azores and Madeira have a generally cooler climate and higher rainfall than the Canaries and Cape Verde.

The islands have a unique biogeography, and are home to several distinct plant and animal communities. None of the Macaronesian islands were part of a continent, so the native plants and animals reached the islands via long-distance dispersal. Laurel-leaved forests, called laurisilva, once covered most of the Azores, Madeira, and parts of the Canaries between 400-1200 m altitude (the eastern Canaries and Cape Verde being too dry). These forests resemble the ancient forests that covered the Mediterranean basin and northwestern Africa before cooling and drying of the ice ages. Trees of the genera Apollonias (Lauraceae), Clethra (Clethraceae), Dracaena (Ruscaceae), Ocotea (Lauraceae), Persea (Lauraceae), and Picconia (Oleaceae), which are found in the Macaronesian laurel forests, are also known from fossils to have lived around the Mediterranean before the ice ages.